Unlocked by

• Occam's Spire
• Grizzlemouth's Spire
• Mathilde's Spire
• Verata's Spire
• Ossa's Spire
• Yassith's Spire
• Pahua's Spire
• Ruka's Spire
• Maklain's Spire
• Tizlak's Spire
• Laranthir's Spire
• Svaard's Spire
• Thackeray's Pillar
• The Twins' Spire
• Zehtuka's Spire
• Nadijeh's Spire
• Tixx's Spire
• Giftbringer's Spire
• Nerashi's Spire
• Togo's Spire